Single-Species
Oncobreds
Arilbreds are often described in terms of aril characteristics attributable to individual aril species, but modern cultivars typically have aril ancestry traceable to more than one species so it's difficult to link visible traits with certain species. One solution is to collect and observe single-species arilbreds. The original first-generation crosses between aril species and TBs provide obvious, easy-to-identify examples of species traits – if you can find them. There are also a number of second-generation crosses that have only one aril species in their ancestry.
The following lists are not comprehensive. They were compiled over 25 years ago for a few species of special interest, drawing on information in the 1976 Checklist.
I.
gatesii
The following varieties came from William Mohr, whose partial fertility became widely known. While imposters have been distributed in the past 30 years, I've found no evidence that labeling errors affected these early introductions. I. gatesii was especially valued for its size and wide form, and if these flowers are compared to TBs of the period, its influence can be readily seen. In the absence of a chromosome count, all are presumed triploids. In theory, any could be used with a modern TB to produce a quarterbred with a full set of . I. gatesii chromosomes. In practice, their fertility is limited and it would probably take many crosses to attain the desired results.
Alta Mohr (Paul, 1948) William Mohr X Alta California.
Amohr (Tompkins, 1946) William Mohr X Sass 40-306.
Barrimohr (Miller, 1940) William Mohr X Unknown
Bella Mohr (Ohl, 1955) William Mohr X Sable
Blumohr (Marx, 1949) William Mohr X Orloff
British Empire (Aylett, NR, introduced in 1934) William Mohr X Pride of Wagga
Casino (Plough, 1953) William Mohr X Firecracker
Colossal (Marx, 1948) William Mohr X El Capitan
Easter Mohr (Weidner, 1949) William Mohr X Easter Morn
Elmohr (Loomis-Long, 1942) William Mohr X Red Seedling
Evermohr (Rakow, 1947) William Mohr X City of Lincoln
Gaymohr (Green, 1957) William Mohr X Winter Carnival
Grace Mohr (Jory, 1935) William Mohr X Unknown TB
Gray Mohr (Muhlestein, 1956) William Mohr X (Alta California x King Midas)
Green Mohr (Muhlestein, 1949) William Mohr X Chosen
Gros Grain (Craig, 1950) William Mohr X (Purissima x Gudrun)
Kulinor (Crawford, 1965) William Mohr X Pink Lace
Laurel Hill (Plough, 1948) William Mohr X (Esplendido x Bruno)
Marbled Mohr (Muhlestein, 1958) William Mohr X ((Midwest x Madame Louis Aureau) x Aldura).
Mardin (Lincoln, 1954) William Mohr X Seedling
Master Neil (Jensen-Muhlestein, 1952) William Mohr X Stained Glass
Mohr Affair (Walker, 1952) William Mohr X Golden Majesty
Mohr Courageous (Naylor, 1953) William Mohr X TB Seedling
Mohrdyke (Aylett, 1946) William Mohr X W. R. Dykes
Mohr Majesty (Walker, 1949) William Mohr X Golden Majesty
Mohrson (White, 1934) William Mohr X Unknown
Mohr Symphony (Gaylon, 1952) William Mohr X Violet Symphony
Mount Mazama (Bailey, 1953) William Mohr X Morocco
My Day (Tharp, 1942) William Mohr X Los Angeles
New Look (Johnson, 1951) William Mohr X Bruno (probably)
N. J. Thomas (Thomas, 1944) William Mohr X El Capitan
Onliwon (Wild, 1954) (W. R. Dykes x Adobe) X William Mohr
Ormohr (Kleinsorge, 1937) William Mohr X Anakim. Counted as a 46-chromosome ABBB-type quarterbred.
Purissamohr (Weidner, 1949) William Mohr X (Purissima x Easter Morn)
Sikh-mohr (Weidner, 1949) William Mohr X (Dauntless x Sikh)
Soquel (Reinelt, 1940) William Mohr X Sunol
Spring Violet (Holmberg, 1959) William Mohr X Ariel
Suez (Schreiner, 1944) William Mohr X Madame Louis Aureau
Vici Lynn Reed (Savage, 1966) Golden Eagle X William Mohr
I.
iberica
Once fully fertile amphidiploids were available, it became possible to produce true chromosome-set quarterbreds by crossing them with tetraploid TBs. The following came from Ib-Mac, any are still worth growing, simply to see examples of aril traits that are inheritable from I. iberica:
Alpine Halo (Brown, 1966) (Green Halo x Snow Elf) X Ib-Mac
Anine (Knopf, 1956) Snow Flurry X Ib-Mac
Argyll Lass (Street, 1963) (Adornment x Sulina) X Ib-Mac
Aril Lady (Danielson, 1975) Katherine Tiffany X Ib-Mac
Beautiful Dreamer (Reynolds, 1965) (Convention Queen x 53-51) X Ib-Mac
Cinna Mac (Hopson, 1956) (Tiffany x White Ruffles) X Ib-Mac
Dance of Spring (Burbridge, 1959) White Ruffles X Ib-Mac
Hidden Secret (Nichols, 1974) Eye Shadow X Ib-Mac
Moonchild (Craig, 1955) Moon Goddess X Ib-Mac
Persian Pattern (Craig, 1950) Gay Senorita X Ib-Mac
Silver Charm (Craig, 1948) Purissima X Ib-Mac
Somewhere (Wall, 1963) Pink Formal X Ib-Mac
I.
paradoxa
Arilarians have long known this species as an interesting novelty, but few have considered its form desirable for hybridizing.
Lady Lilford (Sir Michael Foster, NR, introduced by 1916) I. paradoxa X I. pallida. This was introduced before the registration system was adopted. After it had established its reputation for fertility, it was counted as a 46-chromosome tetraploid. The catch is that this is not a logical composition for its pedigree. If both parents had produced unreduced gametes, the count should be 44 chromosomes. If the reported fertility was actually that of an imposter, the 46-chromosome count can be explained in at least two ways. For example, an unreduced I. paradoxa gamete could have combined with a normal gamete from one of the early 52-chromosome TBs. Or an unreduced gamete of a diploid I. paradoxa offspring could have combined with a normal gamete of a 48-chromosome TB. In any case, both the "Lady Lilford" that was used in hybridizing and the one that was counted probably did come from I. paradoxa because its tendency to produce small falls was well-known and any purported first-generation offspring without this trait would surely have been questioned.
I've been able to identify two quantum-system quarterbreds registered from Lady Lilford. Although I have not seen either of them, if they still exist they should provide insight into the inheritability of traits from I. paradoxa.
Cactiforium (Aylett, NR, introduced 1932) Alcazar X Lady Lilford
Emily Pyke (Aylett, NR, introduced 1933) Kashmir White X Lady Lilford
Diploid offspring of I. paradoxa also provide valuable source material for genetic experiments.
For example:
Par-Pall (Sir Michael Foster, NR) Introduced before the registration system was adopted. This has contributed genes to the modern arilbreds through its offspring, Don Ricardo.
Don Ricardo (Rosenzweig, 1964). Par-Pall X Ib-Mac. I have not yet found a chromosome count, but this is either an unusually fertile triploid or a fully fertile amphidiploid that resulted from an unreduced gamete of the pod parent. Note that this also includes chromosomes from I. iberica, so it's not a single-species oncobred but has been a valuable breeder.
I.
susiana
From Zwanenburg, which was counted as an amphidiploid, there were a number of quarterbreds derived solely from I. susiana. Although this species has a well-deserved reputation for being recessive, it's dotting and veining sometimes show through in its offspring.
Brownie (Douglas, 1954) (Minnie Colquitt x pumila) X Zwanenburg
Buffon (Baker, 1938) BDB X Zwanenburg
Butterfly Wings (White, NR, Introduced 1946) (Pink Jadu X I. susiana) X (Theme x Sacramento)
Califa Dotu (Graves, 1970) Knotty Pine X Zwanenburg
Green Glory (Green, 1939) Leota X Zwanenburg
Mohresque (Keith, 1949) Doa (probably) X Zwanenburg
Mt. Kosciusko (Aylett, 1933) Kashmir White X Zwanenburg
Probably the best-known, as well as the most controversial of these, is Butterfly Wings. C. G. White registered a TB with that name in 1946, but apparently did not introduce it. Lloyd Austin listed a quarterbred, with complete pedigree, under that name during the period in which arilbreds had to be recorded with ASI but not registered with AIS.